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世界上最长的单词意思是什么世界上最长的单词意思是什么？想必会有不少人好奇吧，下面由店铺为大家带来世界上最长的单词意思。
世界上最长的单词意思是Titin(肌联蛋白)Titin(肌联蛋白)是目前已知的最大的蛋白质，它的分子量为2700kDa(25,000多个氨基端)，长度为1μm，约占肌节的一半，是高度弹性分子。
根据IUPAC(国际理论和应用化学联合会)命名规则，该肌联蛋白的名字足足有189819个字母，是名副其实最大的单词。
一个坐在那啥事都不干快速读完也要读3个多少时。
Methionylthreonylthreonylglutaminylarginyltyrosylglutamylseryl leucylphenylalanylalanylglu...以下省略，是不是感受到很大压力呢?除了最长的单词，下面是常见比较长的单词，也十分有意思呢。
常见长单词unenthusiastically adv. 不热肠地;不体贴地共14个字母acquaaintanceship n. 相识，认识;交往关系共16个字母biotransformation n.[生物] 生物转化共17个字母chemiluminescence n.[化学] 化合光;化学反光，在化学反应中，低温反光共17个字母dihydrostreptomycin n.[药学] 二氢链霉素(结核病特效药)共19个字母electroencephalography n.[医学] 脑波纪录仪共22个字母flibbertigibbet n. 轻浮的女子共15个字母great-granddaughter n. 曾孙女;外曾孙女hyperparathyroidism n.[医学] 副甲状腺机能亢进共19个字母internationalization n. 国际化;国际管理化共20个字母Johannisberger n.一种白葡萄酒共14个字母Kaiserslautern n. [地名]凯撒劳坦(德国西部一城市)共14个字母lexicostatistics n.[语言学]词汇统计共16个字母microminiaturization n.[电子学] (电子设备的)超小型化共20个字母noncontradiction n. 无矛盾共16个字母omnidirectional adj.[电学] 全方向的共15个字母parliamentarianism n.[政治学] 议会主义;议会制政体;议会制政府共18个字母quadricentennial n. 四百年纪念(日);四百年祭典共16个字母radiobroadcasting n.[电子学] 无线电广播共18个字母self-actualization n.[心理学] 自我实现共17个字母tetrachloroethylene n.[化学] 四氯乙烯共19个字母unenthusiastically adv. 不热心地;不关心地;冷淡地共18个字母vasoconstriction n.[医学] 血管紧缩;血管缩小well-proportioned adj. 很平衡的;(身体)很均匀的共16个字母xerophthalmia n.[医学]干眼病，结膜干燥症共13个字母yesterdayness n. 最近;昨天性共13个字母Zoroastrianism n. 袄教，拜火教。
冷知识世界上最长的英文单词从前人们广泛使用录音带和录像带时，如果要从头播放，都是左边的轮盘剩余磁带更多，结构上很像一个向右的三角形；播放时，左边轮盘将磁带传送至右边轮盘，方向也是指向右边的。
所以快进是两个向右的三角形（表示加速传送），快退则是方向相反的两个三角形。
为什么播放器中通常用“三角”表示播放其实鱼也是会口渴的！只不过它们饮水的方法和人类有所不同。
德国一研究所指出，由于鱼分为淡水鱼和咸水鱼，因此两种鱼的饮水方法不同。
咸水鱼用吞水的方法得到水分，淡水鱼则经由食物和鱼鳃得到水分。
鱼也会口渴吗20世界上最长的英文单词有189819个英文字母，它是T IT IN （肌联蛋白）的化学名称。
T IT IN 是一种巨型蛋白质，负责肌肉被动弹性。
要把这个单词全部读完需要花费约3.5小时，图片中仅仅是截取了这个单词的一部分，是不是看起来已经让人觉得晕头转向了呢计算机Bu g的由来1945年9月，编译器的发明者格蕾斯·哈珀正带领她的研究小组构造“马克二型”计算机。
突然，计算机死机了。
哈珀在某个出错的继电器上发现了一只被电死的飞蛾，她将这只飞蛾贴到记事本中并标注了一句“第一个发现虫子实例”。
从此，人们就将计算机错误戏称为“B ug”，将发现Bu g并纠正的过程叫做“D eb ug”。
没有首都的国家瑙鲁共和国（简称瑙鲁）是世界上唯一没有首都的国家。
瑙鲁是位于南太平洋密克罗尼西亚群岛的一个岛国，国土面积为21.3平方公里，是世界上第三小的国家。
全国分为14个区，仅在亚伦设有行政管理中心，地方事务则由各政府委员会负责。
身上长有植物的野生动物是什么树懒是唯一身上长有植物（绿藻等）的野生动物，形状略似猴，生活于热带森林中。
它虽然有脚但是却不能走路，靠得是前肢拖动身体前行。
美国动画大片《冰河世纪》里的Sid就是一只树懒！21。
肌联蛋白（Titin）在肌肉收缩中的作用摘要：1954年Hugh Huxley & Jean Hanson提出了肌丝滑行学说，成为了解释肌纤维收缩分子机制的经典学说。
现阶段的研究发现，肌丝滑行学说不能解释向心活动和离心活动的区别，也不能解释后效力量增加的原因，也没包含肌联蛋白（Titin）。
本文通过在 PubMed和CNKI上检索“muscle contraction”“Titin” “肌丝滑行”等关键词，查阅相关文献的方法，对“Titin在肌肉收缩中的作用”进行分析和综述，总结了肌肉收缩过程的研究现状，也对肌丝滑行学说的丰富和完善提出了展望。
关键词：Titin；肌丝滑行；肌肉收缩；1．历史和背景几个世纪以来，肌肉收缩一直令科学家们着迷。
直到19世纪后期，显微镜技术的发展和应用，才对肌肉的基本结构特征有了进一步的认识，如观察到肌肉是规则的横纹肌。
在 20世纪初，一个激活的肌肉被简单地认为是一种新的弹性体（Gasser and Hill，1924）。
在二十世纪五十年代之前，肌肉收缩和产生力量被认为是肌节中间可见的长蛋白链折叠造成的。
这个缩短被认为是由乳酸引起的。
在五十年代初，仔细分析A带的长度，发现肌球蛋白在各种收缩的条件下都不发生长度变化，因此，不能说明肌肉收缩产生力量的过程中肌肉组织可以经历的长度变化（Huxley，1953）。
1957年，安德鲁?赫胥黎提出了这种相对滑动可能发生的设想，并提供了一个现在被称为肌丝滑行学说的数学框架（Huxley，1957）。
这张纸，被引用超过3000次（Google Scholar，June2014），大致描述了今天我们对肌肉收缩的思考。
2. 对肌丝滑行学说的讨论分析目前有关肌肉如何收缩的模型是肌丝滑动理论，但该理论并没有充分包含巨型肌联蛋白、肌肉中固有的弹性能力，也没有肌肉离心和向心活动的区别，同样也没有涉及后效力量的增加。
肌丝滑行学说认为肌肉的缩短是由于肌小节中细肌丝在粗肌丝之间滑行造成的，即当肌肉收缩时由Z线发出的细肌丝在某种力量的作用下向A带中央滑行，结果相邻的各Z线互相靠近，肌小节的长度变短，从而导致肌纤维的缩短。
这是⼀个包括189819个字母的肌联蛋⽩的化学名1. Methionylalanylthreonylserylarginylglycylalanylserylarginylcysteinylproly-2. larginylaspartylisoleucylalanylasparaginylvalylmethionylglutaminylarginyl-3. leucylglutaminylaspartylglutamylglutaminylglutamylisoleucylvalylglutaminy-4. llysylarginylthreonylphenylalanylthreonyllysyltryptophylisoleucylasparagi-5. nylserylhistidylleucylalanyllysylarginyllysylprolylprolylmethionylvalylva-6. lylaspartylaspartylleucylphenylalanylglutamylaspartylmethionyllysylaspart-7. ylglycylvalyllysylleucylleucylalanylleucylleucylglutamylvalylleucylserylg-8. lycylglutaminyllysylleucylprolylcysteinylglutamylglutaminylglycylarginyla-9. rginylmethionyllysylarginylisoleucylhistidylalanylvalylalanylasparaginyli-10. soleucylglycylthreonylalanylleucyllysylphenylalanylleucylglutamylglycylar-11. ginyllysylisoleucyllysylleucylvalylasparaginylisoleucylasparaginylserylth-12. reonylaspartylisoleucylalanylaspartylglycylarginylprolylserylisoleucylval-13. ylleucylglycylleucylmethionyltryptophylthreonylisoleucylisoleucylleucylty-14. rosylphenylalanylglutaminylisoleucylglutamylglutamylleucylthreonylserylas-15. paraginylleucylprolylglutaminylleucylglutaminylserylleucylserylserylseryl-16. alanylserylserylvalylaspartylserylisoleucylvalylserylserylglutamylthreony-17. lprolylserylprolylprolylseryllysylarginyllysylvalylthreonylthreonyllysyli-18. soleucylglutaminylglycylasparaginylalanyllysyllysylalanylleucylleucyllysy-19. ltryptophylvalylglutaminyltyrosylthreonylalanylglycyllysylglutaminylthreo-20. nylglycylisoleucylglutamylvalyllysylaspartylphenylalanylglycyllysylserylt-21. ryptophylarginylserylglycylvalylalanylphenylalanylhistidylserylvalylisole-22. ucylhistidylalanylisoleucylarginylprolylglutamylleucylvalylaspartylleucyl-23. glutamylthreonylvalyllysylglycylarginylserylasparaginylarginylglutamylasp-24. araginylleucylglutamylaspartylalanylphenylalanylthreonylisoleucylalanylgl-25. utamylthreonylglutamylleucylglycylisoleucylprolylarginylleucylleucylaspar-26. tylprolylglutamylaspartylvalylaspartylvalylaspartyllysylprolylaspartylglu-27. tamyllysylserylisoleucylmethionylthreonyltyrosylvalylalanylglutaminylphen-28. ylalanylleucyllysylhistidyltyrosylprolylaspartylisoleucylhistidylasparagi-29. nylalanylserylthreonylaspartylglycylglutaminylglutamylaspartylaspartylglu-30. tamylisoleucylleucylprolylglycylphenylalanylprolylserylphenylalanylalanyl-31. asparaginylserylvalylglutaminylasparaginylphenylalanyllysylarginylglutamy-32. laspartylarginylvalylisoleucylphenylalanyllysylglutamylmethionyllysylvaly-33. ltryptophylisoleucylglutamylglutaminylphenylalanylglutamylarginylaspartyl-34. leucylthreonylarginylalanylglutaminylmethionylvalylglutamylserylasparagin-35. ylleucylglutaminylaspartyllysyltyrosylglutaminylserylphenylalanyllysylhis-36. tidylphenylalanylarginylvalylglutaminyltyrosylglutamylmethionyllysylargin-37. yllysylglutaminylisoleucylglutamylhistidylleucylisoleucylglutaminylprolyl-38. leucylhistidylarginylaspartylglycyllysylleucylserylleucylaspartylglutamin-39. ylalanylleucylvalyllysylglutaminylseryltryptophylaspartylarginylvalylthre-40. onylserylarginylleucylphenylalanylaspartyltryptophylhistidylisoleucylglut-41. aminylleucylaspartyllysylserylleucylprolylalanylprolylleucylglycylthreony-42. lisoleucylglycylalanyltryptophylleucyltyrosylarginylalanylglutamylvalylal-43. anylleucylarginylglutamylglutamylisoleucylthreonylvalylglutaminylglutamin-44. ylvalylhistidylglutamylglutamylthreonylalanylasparaginylthreonylisoleucyl-45. glutaminylarginyllysylleucylglutamylglutaminylhistidyllysylaspartylleucyl-46. leucylglutaminylasparaginylthreonylaspartylalanylhistidyllysylarginylalan-47. ylphenylalanylhistidylglutamylisoleucyltyrosylarginylthreonylarginylseryl-48. valylasparaginylglycylisoleucylprolylvalylprolylprolylaspartylglutaminyll-49. eucylglutamylaspartylmethionylalanylglutamylarginylphenylalanylhistidylph-50. enylalanylvalylserylserylthreonylserylglutamylleucylhistidylleucylmethion-51. yllysylmethionylglutamylphenylalanylleucylglutamylleucyllysyltyrosylargin-52. ylleucylleucylserylleucylleucylvalylleucylalanylglutamylseryllysylleucyll-53. ysylseryltryptophylisoleucylisoleucyllysyltyrosylglycylarginylarginylglut-54. amylserylvalylglutamylglutaminylleucylleucylglutaminylasparaginyltyrosylv-55. alylserylphenylalanylisoleucylglutamylasparaginylseryllysylphenylalanylph-56. enylalanylglutamylglutaminyltyrosylglutamylvalylthreonyltyrosylglutaminyl-57. isoleucylleucyllysylglutaminylthreonylalanylglutamylmethionyltyrosylvalyl-58. lysylalanylaspartylglycylserylvalylglutamylglutamylalanylglutamylasparagi-59. nylvalylmethionyllysylphenylalanylmethionylasparaginylglutamylthreonylthr-60. eonylalanylglutaminyltryptophylarginylasparaginylleucylserylvalylglutamyl-61. valylarginylserylvalylarginylserylmethionylleucylglutamylglutamylvalyliso-62. leucylserylasparaginyltryptophylaspartylarginyltyrosylglycylasparaginylth-63. reonylvalylalanylserylleucylglutaminylalanyltryptophylleucylglutamylaspar-64. tylalanylglutamyllysylmethionylleucylasparaginylglutaminylserylglutamylas-65. paraginylalanyllysyllysylaspartylphenylalanylphenylalanylarginylasparagin-66. ylleucylprolylhistidyltryptophylisoleucylglutaminylglutaminylhistidylthre-67. onylalanylmethionylasparaginylaspartylalanylglycylasparaginylphenylalanyl-68. leucylisoleucylglutamylthreonylcysteinylaspartylglutamylmethionylvalylser-69. ylarginylaspartylleucyllysylglutaminylglutaminylleucylleucylleucylleucyla-70. sparaginylglycylarginyltryptophylarginylglutamylleucylphenylalanylmethion-71. ylglutamylvalyllysylglutaminyltyrosylalanylglutaminylalanylaspartylglutam-72. ylmethionylaspartylarginylmethionyllysyllysylglutamyltyrosylthreonylaspar-73. tylcysteinylvalylvalylthreonylleucylserylalanylphenylalanylalanylthreonyl-74. glutamylalanylhistidyllysyllysylleucylserylglutamylprolylleucylglutamylva-75. lylserylphenylalanylmethionylasparaginylvalyllysylleucylleucylisoleucylgl-76. utaminylaspartylleucylglutamylaspartylisoleucylglutamylglutaminylarginylv-77. alylprolylvalylmethionylaspartylalanylglutaminyltyrosyllysylisoleucylisol-78. eucylthreonyllysylthreonylalanylhistidylleucylisoleucylthreonyllysylgluta-79. mylserylprolylglutaminylglutamylglutamylglycyllysylglutamylmethionylpheny-80. lalanylalanylthreonylmethionylseryllysylleucyllysylglutamylglutaminylleuc-81. ylthreonyllysylvalyllysylglutamylcysteinyltyrosylserylprolylleucylleucylt-82. yrosylglutamylserylglutaminylglutaminylleucylleucylisoleucylprolylleucylg-83. lutamylglutamylleucylglutamyllysylglutaminylmethionylthreonylserylphenyla-84. lanyltyrosylaspartylserylleucylglycyllysylisoleucylasparaginylglutamyliso-85. leucylisoleucylthreonylvalylleucylglutamylarginylglutamylalanylglutaminyl-86. serylserylalanylleucylphenylalanyllysylglutaminyllysylhistidylglutaminylg-87. lutamylleucylleucylalanylcysteinylglutaminylglutamylasparaginylcysteinyll-88. ysyllysylthreonylleucylthreonylleucylisoleucylglutamyllysylglycylserylglu-89. taminylserylvalylglutaminyllysylphenylalanylvalylthreonylleucylserylaspar-90. aginylvalylleucyllysylhistidylphenylalanylaspartylglutaminylthreonylargin-91. ylleucylglutaminylarginylglutaminylisoleucylalanylaspartylisoleucylhistid-92. ylvalylalanylphenylalanylglutaminylserylmethionylvalyllysyllysylthreonylg-93. lycylaspartyltryptophyllysyllysylhistidylvalylglutamylthreonylasparaginyl-94. serylarginylleucylmethionyllysyllysylphenylalanylglutamylglutamylserylarg-95. inylalanylglutamylleucylglutamyllysylvalylleucylarginylisoleucylalanylglu-96. taminylglutamylglycylleucylglutamylglutamyllysylglycylaspartylprolylgluta-97. mylglutamylleucylleucylarginylarginylhistidylthreonylglutamylphenylalanyl-98. phenylalanylserylglutaminylleucylaspartylglutaminylarginylvalylleucylaspa-99. raginylalanylphenylalanylleucyllysylalanylcysteinylaspartylglutamylleucyl-100. threonylaspartylisoleucylleucylprolylglutamylglutaminylglutamylglutaminyl-101. glutaminylglycylleucylglutaminylglutamylalanylvalylarginyllysylleucylhist-102. idyllysylglutaminyltryptophyllysylaspartylleucylglutaminylglycylglutamyla-103. lanylprolyltyrosylhistidylleucylleucylhistidylleucyllysylisoleucylasparty-104. lvalylglutamyllysylasparaginylarginylphenylalanylleucylalanylserylvalylgl-105. utamylglutamylcysteinylarginylthreonylglutamylleucylaspartylarginylglutam-106. ylthreonyllysylleucylmethionylprolylglutaminylglutamylglycylserylglutamyl-107. lysylisoleucylisoleucyllysylglutamylhistidylarginylvalylphenylalanylpheny-108. lalanylserylaspartyllysylglycylprolylhistidylhistidylleucylcysteinylgluta-109. myllysylarginylleucylglutaminylleucylisoleucylglutamylglutamylleucylcyste-110. inylvalyllysylleucylprolylvalylarginylaspartylprolylvalylarginylaspartylt-111. hreonylprolylglycylthreonylcysteinylhistidylvalylthreonylleucyllysylgluta-112. mylleucylarginylalanylalanylisoleucylaspartylserylthreonyltyrosylarginyll-113. ysylleucylmethionylglutamylaspartylprolylaspartyllysyltryptophyllysylaspa-114. rtyltyrosylthreonylserylarginylphenylalanylserylglutamylphenylalanylseryl-115. seryltryptophylisoleucylserylthreonylasparaginylglutamylthreonylglutaminy-116. lleucyllysylglycylisoleucyllysylglycylglutamylalanylisoleucylaspartylthre-117. onylalanylasparaginylhistidylglycylglutamylvalyllysylarginylalanylvalylgl-118. utamylglutamylisoleucylarginylasparaginylglycylvalylthreonyllysylarginylg-119. lycylglutamylthreonylleucylseryltryptophylleucyllysylserylarginylleucylly-120. sylvalylleucylthreonylglutamylvalylserylserylglutamylasparaginylglutamyla-121. lanylglutaminyllysylglutaminylglycylaspartylglutamylleucylalanyllysylleuc-122. ylserylserylserylphenylalanyllysylalanylleucylvalylthreonylleucylleucylse-123. rylglutamylvalylglutamyllysylmethionylleucylserylasparaginylphenylalanylg-124. lycylaspartylcysteinylvalylglutaminyltyrosyllysylglutamylisoleucylvalylly-125. sylasparaginylserylleucylglutamylglutamylleucylisoleucylserylglycylseryll-126. ysylglutamylvalylglutaminylglutamylglutaminylalanylglutamyllysylisoleucyl-127. leucylaspartylthreonylglutamylasparaginylleucylphenylalanylglutamylalanyl-128. glutaminylglutaminylleucylleucylleucylhistidylhistidylglutaminylglutaminy-129. llysylthreonyllysylarginylisoleucylserylalanyllysyllysylarginylaspartylva-130. lylglutaminylglutaminylglutaminylisoleucylalanylglutaminylalanylglutaminy-131. lglutaminylglycylglutamylglycylglycylleucylprolylaspartylarginylglycylhis-132. tidylglutamylglutamylleucylarginyllysylleucylglutamylserylthreonylleucyla-133. spartylglycylleucylglutamylarginylserylarginylglutamylarginylglutaminylgl-134. utamylarginylarginylisoleucylglutaminylvalylthreonylleucylarginyllysyltry-135. ptophylglutamylarginylphenylalanylglutamylthreonylasparaginyllysylglutamy-136. lthreonylvalylvalylarginyltyrosylleucylphenylalanylglutaminylthreonylglyc-137. ylserylserylhistidylglutamylarginylphenylalanylleucylserylphenylalanylser-138. ylserylleucylglutamylserylleucylserylserylglutamylleucylglutamylglutaminy-139. lthreonyllysylglutamylphenylalanylseryllysylarginylthreonylglutamylseryli-140. soleucylalanylvalylglutaminylalanylglutamylasparaginylleucylvalyllysylglu-141. tamylalanylserylglutamylisoleucylprolylleucylglycylprolylglutaminylaspara-142. ginyllysylglutaminylleucylleucylglutaminylglutaminylglutaminylalanyllysyl-143. serylisoleucyllysylglutamylglutaminylvalyllysyllysylleucylglutamylasparty-144. lthreonylleucylglutamylglutamylaspartylisoleucyllysylthreonylmethionylglu-145. tamylmethionylvalyllysylthreonyllysyltryptophylaspartylhistidylphenylalan-146. ylglycylserylasparaginylphenylalanylglutamylthreonylleucylserylvalyltrypt-147. ophylisoleucylthreonylglutamyllysylglutamyllysylglutamylleucylasparaginyl-148. alanylleucylglutamylthreonylserylserylserylalanylmethionylaspartylmethion-149. ylglutaminylisoleucylserylglutaminylisoleucyllysylvalylthreonylisoleucylg-150. lutaminylglutamylisoleucylglutamylseryllysylleucylserylserylisoleucylvaly-151. lglycylleucylglutamylglutamylglutamylalanylglutaminylserylphenylalanylala-152. nylglutaminylphenylalanylvalylthreonylthreonylglycylglutamylserylalanylar-153. ginylisoleucyllysylalanyllysylleucylthreonylglutaminylisoleucylarginylarg-154. inyltyrosylglycylglutamylglutamylleucylarginylglutamylhistidylalanylgluta-155. minylcysteinylleucylglutamylglycylthreonylisoleucylleucylglycylhistidylle-156. ucylserylglutaminylglutaminylglutaminyllysylphenylalanylglutamylglutamyla-157. sparaginylleucylarginyllysylisoleucylglutaminylglutaminylserylvalylserylg-158. lutamylphenylalanylglutamylaspartyllysylleucylalanylvalylprolylisoleucyll-159. ysylisoleucylcysteinylserylserylalanylthreonylglutamylthreonyltyrosyllysy-160. lvalylleucylglutaminylglutamylhistidylmethionylaspartylleucylcysteinylglu-161. taminylalanylleucylglutamylserylleucylserylserylalanylisoleucylthreonylal-162. anylphenylalanylserylalanylserylalanylarginyllysylvalylvalylasparaginylar-163. ginylaspartylserylcysteinylvalylglutaminylglutamylalanylalanylalanylleucy-164. lglutaminylglutaminylglutaminyltyrosylglutamylaspartylisoleucylleucylargi-165. nylarginylalanyllysylglutamylarginylglutaminylthreonylalanylleucylglutamy-166. lasparaginylleucylleucylalanylhistidyltryptophylglutaminylarginylleucylgl-167. utamyllysylglutamylleucylserylserylphenylalanylleucylthreonyltryptophylle-168. ucylglutamylarginylglycylglutamylalanyllysylalanylserylserylprolylglutamy-169. lmethionylaspartylisoleucylserylalanylaspartylarginylvalyllysylvalylgluta-170. mylglycylglutamylleucylglutaminylleucylisoleucylglutaminylalanylleucylglu-171. taminylasparaginylglutamylvalylvalylserylglutaminylalanylserylphenylalany-172. ltyrosylseryllysylleucylleucylglutaminylleucyllysylglutamylserylleucylphe-173. nylalanylserylvalylalanylseryllysylaspartylaspartylvalyllysylmethionylmet-174. hionyllysylleucylhistidylleucylglutamylglutaminylleucylaspartylglutamylar-175. ginyltryptophylarginylaspartylleucylprolylglutaminylisoleucylisoleucylasp-176. araginyllysylarginylisoleucylasparaginylphenylalanylleucylglutaminylseryl-177. valylvalylalanylglutamylhistidylglutaminylglutaminylphenylalanylaspartylg-178. lutamylleucylleucylleucylserylphenylalanylserylvalyltryptophylisoleucylly-179. sylleucylphenylalanylleucylserylglutamylleucylglutaminylthreonylthreonyls-180. erylglutamylisoleucylserylisoleucylmethionylaspartylhistidylglutaminylval-181. ylalanylleucylthreonylarginylhistidyllysylaspartylhistidylalanylalanylglu-182. tamylvalylglutamylseryllysyllysylglycylglutamylleucylglutaminylserylleucy-183. lglutaminylglycylhistidylleucylalanyllysylleucylglycylserylleucylglycylar-184. ginylalanylglutamylaspartylleucylhistidylleucylleucylglutaminylglycyllysy-185. lalanylglutamylaspartylcysteinylphenylalanylglutaminylleucylphenylalanylg-186. lutamylglutamylalanylserylglutaminylvalylvalylglutamylarginylarginylgluta-187. minylleucylalanylleucylserylhistidylleucylalanylglutamylphenylalanylleucy-188. lglutaminylserylhistidylalanylserylleucylserylglycylisoleucylleucylarginy-189. lglutaminylleucylarginylglutaminylthreonylvalylglutamylalanylthreonylaspa-190. raginylserylmethionylasparaginyllysylasparaginylglutamylserylaspartylleuc-191. ylisoleucylglutamyllysylaspartylleucylasparaginylaspartylalanylleucylglut-192. aminylasparaginylalanyllysylalanylleucylglutamylserylalanylalanylvalylser-193. ylleucylaspartylglycylisoleucylleucylseryllysylalanylglutaminyltyrosylhis-194. tidylleucyllysylisoleucylglycylserylserylglutamylglutaminylarginylthreony-195. lserylcysteinylarginylalanylthreonylalanylaspartylglutaminylleucylcystein-196. ylglycylglutamylvalylglutamylarginylisoleucylglutaminylasparaginylleucyll-197. eucylglycylthreonyllysylglutaminylserylglutamylalanylaspartylalanylleucyl-198. alanylvalylleucyllysyllysylalanylphenylalanylglutaminylaspartylglutaminyl-199. lysylglutamylglutamylleucylleucyllysylserylisoleucylglutamylaspartylisole-200. ucylglutamylglutamylarginylthreonylaspartyllysylglutamylarginylleucyllysy-201. lglutamylprolylthreonylarginylglutaminylalanylleucylglutaminylglutaminyla-202. rginylleucylarginylvalylphenylalanylasparaginylglutaminylleucylglutamylas-203. partylglutamylleucylasparaginylserylhistidylglutamylhistidylglutamylleucy-204. lcysteinyltryptophylleucyllysylaspartyllysylalanyllysylglutaminylisoleucy-205. lalanylglutaminyllysylaspartylvalylalanylphenylalanylalanylprolylglutamyl-206. valylaspartylarginylglutamylisoleucylasparaginylarginylleucylglutamylvaly-207. lthreonyltryptophylaspartylaspartylthreonyllysylarginylleucylisoleucylhis-208. tidylglutamylasparaginylglutaminylglycylglutaminylcysteinylcysteinylglycy-209. lleucylisoleucylaspartylleucylmethionylarginylglutamyltyrosylglutaminylas-210. paraginylleucyllysylserylalanylvalylseryllysylvalylleucylglutamylasparagi-211. nylalanylserylserylvalylisoleucylvalylthreonylarginylthreonylthreonylisol-212. eucyllysylaspartylglutaminylglutamylaspartylleucyllysyltryptophylalanylph-213. enylalanylseryllysylhistidylglutamylthreonylalanyllysylasparaginyllysylme-214. thionylasparaginyltyrosyllysylglutaminyllysylaspartylleucylaspartylaspara-215. ginylphenylalanylthreonylseryllysylglycyllysylhistidylleucylleucylserylgl-216. utamylleucyllysyllysylisoleucylhistidylserylserylaspartylphenylalanylsery-217. lleucylvalyllysylthreonylaspartylmethionylglutamylserylthreonylvalylaspar-218. tyllysyltryptophylleucylaspartylvalylserylglutamyllysylleucylglutamylglut-219. amylasparaginylmethionylaspartylarginylleucylarginylvalylserylleucylseryl-220. isoleucyltryptophylaspartylaspartylvalylleucylserylthreonylarginylasparty-221. lglutamylisoleucylglutamylglycyltryptophylserylasparaginylasparaginylcyst-222. einylvalylprolylglutaminylmethionylalanylglutamylasparaginylisoleucylsery-223. lasparaginylleucylaspartylasparaginylhistidylleucylarginylalanylglutamylg-224. lutamylleucylleucyllysylglutamylphenylalanylglutamylserylglutamylvalyllys-225. ylasparaginyllysylalanylleucylarginylleucylglutamylglutamylleucylhistidyl-226. seryllysylvalylasparaginylaspartylleucyllysylglutamylleucylthreonyllysyla-227. sparaginylleucylglutamylthreonylprolylprolylaspartylleucylglutaminylpheny-228. lalanylisoleucylglutamylalanylaspartylleucylmethionylglutaminyllysylleucy-229. lglutamylhistidylalanyllysylglutamylisoleucylthreonylglutamylvalylalanyll-230. ysylglycylthreonylleucyllysylaspartylphenylalanylthreonylalanylglutaminyl-231. serylthreonylglutaminylvalylglutamyllysylphenylalanylisoleucylasparaginyl-232. aspartylisoleucylthreonylthreonyltryptophylphenylalanylthreonyllysylvalyl-233. glutamylglutamylserylleucylmethionylasparaginylcysteinylalanylglutaminyla-234. sparaginylglutamylthreonylcysteinylglutamylalanylleucyllysyllysylvalyllys-235. ylaspartylisoleucylglutaminyllysylglutamylleucylglutaminylserylglutaminyl-236. glutaminylserylasparaginylisoleucylserylserylthreonylglutaminylglutamylas-237. paraginylleucylasparaginylserylleucylcysteinylarginyllysyltyrosylhistidyl-238. serylalanylglutamylleucylglutamylserylleucylglycylarginylalanylmethionylt-239. hreonylglycylleucylisoleucyllysyllysylhistidylglutamylalanylvalylserylglu-240. taminylleucylcysteinylseryllysylthreonylglutaminylalanylserylleucylglutam-241. inylglutamylserylleucylglutamyllysylhistidylphenylalanylserylglutamylsery-242. lmethionylglutaminylglutamylphenylalanylglutaminylglutamyltryptophylpheny-243. lalanylleucylglycylalanyllysylalanylalanylalanyllysylglutamylserylserylas-244. partylarginylthreonylglycylaspartylseryllysylvalylleucylglutamylalanyllys-245. ylleucylhistidylaspartylleucylglutaminylasparaginylisoleucylleucylasparty-246. lserylvalylserylaspartylglycylglutaminylseryllysylleucylaspartylalanylval-247. ylthreonylglutaminylglutamylglycylglutaminylthreonylleucyltyrosylalanylhi-248. stidylleucylseryllysylglutaminylisoleucylvalylserylserylisoleucylglutamin-249. ylglutamylglutaminylisoleucylthreonyllysylalanylasparaginylglutamylglutam-250. ylphenylalanylglutaminylalanylphenylalanylleucyllysylglutaminylcysteinyll-251. eucyllysylaspartyllysylglutaminylalanylleucylglutaminylaspartylcysteinyla-252. lanylserylglutamylleucylglycylserylphenylalanylglutamylaspartylglutaminyl-253. histidylarginyllysylleucylasparaginylleucyltryptophylisoleucylhistidylglu-254. tamylmethionylglutamylglutamylarginylphenylalanylasparaginylthreonylgluta-255. mylasparaginylleucylglycylglutamylseryllysylglutaminylhistidylisoleucylpr-256. olylglutamyllysyllysylasparaginylglutamylvalylhistidyllysylvalylglutamylm-257. ethionylphenylalanylleucylglycylglutamylleucylleucylalanylalanylarginylgl-258. utamylserylleucylaspartyllysylleucylserylglutaminylarginylglycylglutaminy-259. lleucylleucylserylglutamylglutamylglycylhistidylglycylalanylglycylglutami-260. nylglutamylglycylarginylleucylcysteinylserylglutaminylleucylleucylthreony-261. lserylhistidylglutaminylasparaginylleucylleucylarginylmethionylthreonylly-262. sylglutamyllysylleucylarginylserylcysteinylglutaminylvalylalanylleucylglu-263. taminylglutamylhistidylglutamylalanylleucylglutamylglutamylalanylleucylgl-264. utaminylserylmethionyltryptophylphenylalanyltryptophylvalyllysylalanyliso-265. leucylglutaminylaspartylarginylleucylalanylcysteinylalanylglutamylserylth-266. reonylleucylglycylseryllysylaspartylthreonylleucylglutamyllysylarginylleu-267. cylserylglutaminylisoleucylglutaminylaspartylisoleucylleucylleucylmethion-268. yllysylglycylglutamylglycylglutamylvalyllysylleucylasparaginylmethionylal-269. anylisoleucylglycyllysylglycylglutamylglutaminylalanylleucylarginylseryls-270. erylasparaginyllysylglutamylglycylglutaminylarginylvalylisoleucylglutamin-271. ylthreonylglutaminylleucylglutamylthreonylleucyllysylglutamylvalyltryptop-272. hylalanylaspartylisoleucylmethionylserylserylserylvalylhistidylalanylglut-273. aminylserylthreonylleucylglutamylserylvalylisoleucylserylglutaminyltrypto-274. phylasparaginylaspartyltyrosylvalylglutamylarginyllysylasparaginylglutami-275. nylleucylglutamylglutaminyltryptophylmethionylglutamylserylvalylaspartylg-276. lutaminyllysylisoleucylglutamylhistidylprolylleucylglutaminylprolylglutam-277. inylprolylglycylleucyllysylglutamyllysylphenylalanylvalylleucylleucylaspa-278. rtylhistidylleucylglutaminylserylisoleucylleucylserylglutamylalanylglutam-279. ylaspartylhistidylthreonylarginylalanylleucylhistidylarginylleucylisoleuc-280. ylalanyllysylserylarginylglutamylleucyltyrosylglutamyllysylthreonylglutam-281. ylaspartylglutamylserylphenylalanyllysylaspartylthreonylalanylglutaminylg-282. lutamylglutamylleucyllysylthreonylglutaminylphenylalanylasparaginylaspart-283. ylisoleucylmethionylthreonylvalylalanyllysylglutamyllysylmethionylarginyl-284. lysylvalylglutamylglutamylisoleucylvalyllysylaspartylhistidylleucylmethio-285. nyltyrosylleucylaspartylalanylvalylhistidylglutamylphenylalanylthreonylas-286. partyltryptophylleucylhistidylserylalanyllysylglutamylglutamylleucylhisti-287. dylarginyltryptophylserylaspartylmethionylserylglycylaspartylserylserylal-288. anylthreonylglutaminyllysyllysylleucylseryllysylisoleucyllysylglutamylleu-289. cylisoleucylaspartylserylarginylglutamylisoleucylglycylalanylserylarginyl-290. leucylserylarginylvalylglutamylserylleucylalanylprolylglutamylvalyllysylg-291. lutaminylasparaginylthreonylthreonylalanylserylglycylcysteinylglutamylleu-292. cylmethionylhistidylthreonylglutamylmethionylglutaminylalanylleucylarginy-293. lalanylaspartyltryptophyllysylglutaminyltryptophylglutamylaspartylserylva-294. lylphenylalanylglutaminylthreonylglutaminylserylcysteinylleucylglutamylas-295. paraginylleucylvalylserylglutaminylmethionylalanylleucylserylglutamylglut-296. aminylglutamylphenylalanylserylglycylglutaminylvalylalanylglutaminylleucy-297. lglutamylglutaminylalanylleucylglutamylglutaminylphenylalanylserylalanyll-298. eucylleucyllysylthreonyltryptophylalanylglutaminylglutaminylleucylthreony-299. lleucylleucylglutamylglycyllysylasparaginylthreonylaspartylglutamylglutam-300. ylisoleucylvalylglutamylcysteinyltryptophylhistidyllysylglycylglutaminylg-301. lutamylisoleucylleucylaspartylalanylleucylglutaminyllysylalanylglutamylpr-302. olylarginylthreonylglutamylaspartylleucyllysylserylglutaminylleucylaspara-303. ginylglutamylleucylcysteinylarginylphenylalanylserylarginylaspartylleucyl-304. serylthreonyltyrosylserylglycyllysylvalylserylglycylleucylisoleucyllysylg-305. lutamyltyrosylasparaginylcysteinylleucylcysteinylleucylglutaminylalanylse-306. ryllysylglycylcysteinylglutaminylasparaginyllysylglutamylglutaminylisoleu-307. cylleucylglutaminylglutaminylarginylphenylalanylarginyllysylalanylphenyla-308. lanylarginylaspartylphenylalanylglutaminylglutaminyltryptophylleucylvalyl-309. asparaginylalanyllysylisoleucylthreonylthreonylalanyllysylcysteinylphenyl-310. alanylaspartylisoleucylprolylglutaminylasparaginylisoleucylserylglutamylv-311. alylserylthreonylserylleucylglutaminyllysylisoleucylglutaminylglutamylphe-312. nylalanylleucylserylglutamylserylglutamylasparaginylglycylglutaminylhisti-313. dyllysylleucylasparaginylmethionylmethionylleucylseryllysylglycylglutamyl-314. leucylleucylserylthreonylleucylleucylthreonyllysylglutamyllysylalanyllysy-315. lglycylisoleucylglutaminylalanyllysylvalylthreonylalanylalanyllysylglutam-316. ylaspartyltryptophyllysylasparaginylphenylalanylhistidylserylasparaginyll-317. eucylhistidylglutaminyllysylglutamylserylalanylleucylglutamylasparaginyll-318. eucyllysylisoleucylglutaminylmethionyllysylaspartylphenylalanylglutamylva-319. lylserylalanylglutamylprolylisoleucylglutaminylaspartyltryptophylleucylse-320. ryllysylthreonylglutamyllysylmethionylvalylhistidylglutamylserylserylaspa-321. raginylarginylleucyltyrosylaspartylleucylprolylalanyllysylarginylarginylg-322. lutamylglutaminylglutaminyllysylleucylglutaminylserylvalylleucylglutamylg-323. lutamylisoleucylhistidylcysteinyltyrosylglutamylprolylglutaminylleucylasp-324. araginylarginylleucyllysylglutamyllysylalanylglutaminylglutaminylleucyltr-325. yptophylglutamylglycylglutaminylalanylalanylseryllysylserylphenylalanylar-326. ginylhistidylarginylvalylserylglutaminylleucylserylserylglutaminyltyrosyl-327. leucylalanylleucylserylasparaginylleucylthreonyllysylglutamyllysylvalylse-328. rylarginylleucylaspartylarginylisoleucylvalylalanylglutamylhistidylaspara-329. ginylglutaminylphenylalanylserylleucylglycylisoleucyllysylglutamylleucylg-330. lutaminylaspartyltryptophylmethionylthreonylaspartylalanylisoleucylhistid-331. ylmethionylleucylaspartylseryltyrosylcysteinylhistidylprolylthreonylseryl-332. aspartyllysylserylvalylleucylaspartylserylarginylthreonylleucyllysylleucy-333. lglutamylalanylleucylleucylserylvalyllysylglutaminylglutamyllysylglutamyl-334. isoleucylglutaminylmethionyllysylmethionylisoleucylvalylthreonylarginylgl-335. ycylglutamylserylvalylleucylglutaminylasparaginylthreonylserylprolylgluta-336. mylglycylisoleucylprolylthreonylisoleucylglutaminylglutaminylglutaminylle-337. ucylglutaminylserylvalyllysylaspartylmethionyltryptophylalanylserylleucyl-338. leucylserylalanylglycylisoleucylarginylcysteinyllysylserylglutaminylleucy-339. lglutamylglycylalanylleucylseryllysyltryptophylthreonylseryltyrosylglutam-340. inylaspartylglycylvalylarginylglutaminylphenylalanylserylglycyltryptophyl-341. methionylaspartylserylmethionylglutamylalanylasparaginylleucylasparaginyl-342. glutamylserylglutamylarginylglutaminylhistidylalanylglutamylleucylarginyl-343. aspartyllysylthreonylthreonylmethionylleucylglycyllysylalanyllysylleucyll-344. eucylasparaginylglutamylglutamylvalylleucylseryltyrosylserylserylleucylle-345. ucylglutamylthreonylisoleucylglutamylvalyllysylglycylalanylglycylmethiony-346. lthreonylglutamylhistidyltyrosylvalylthreonylglutaminylleucylglutamylleuc-347. ylglutaminylaspartylleucylglutaminylglutamylarginyltyrosylarginylalanylis-348. oleucylglutaminylglutamylarginylalanyllysylglutamylalanylvalylthreonyllys-349. ylserylglutamyllysylleucylvalylarginylleucylhistidylglutaminylglutamyltyr-350. osylglutaminylarginylaspartylleucyllysylalanylphenylalanylglutamylvalyltr-351. yptophylleucylglycylglutaminylglutamylglutaminylglutamyllysylleucylaspart-352. ylglutaminyltyrosylserylvalylleucylglutamylglycylaspartylalanylhistidylth-353. reonylhistidylglutamylthreonylthreonylleucylarginylaspartylleucylglutamin-354. ylglutamylleucylglutaminylvalylhistidylcysteinylalanylglutamylglycylgluta-355. minylalanylleucylleucylasparaginylserylvalylleucylhistidylthreonylarginyl-356. glutamylaspartylvalylisoleucylprolylserylglycylisoleucylprolylglutaminyla-357. lanylglutamylaspartylarginylalanylleucylglutamylserylleucylarginylglutami-358. nylaspartyltryptophylglutaminylalanyltyrosylglutaminylhistidylarginylleuc-359. ylserylglutamylthreonylarginylthreonylglutaminylphenylalanylasparaginylas-360. paraginylvalylvalylasparaginyllysylleucylarginylleucylmethionylglutamylgl-361. utaminyllysylphenylalanylglutaminylglutaminylvalylaspartylglutamyltryptop-362. hylleucyllysylthreonylalanylglutamylglutamyllysylvalylserylprolylarginylt-363. hreonylarginylarginylglutaminylserylasparaginylarginylalanylthreonyllysyl-364. glutamylisoleucylglutaminylleucylhistidylglutaminylmethionyllysyllysyltry-365. ptophylhistidylglutamylglutamylvalylthreonylalanyltyrosylarginylaspartylg-366. lutamylvalylglutamylglutamylvalylglycylalanylarginylalanylglutaminylgluta-367. mylisoleucylleucylaspartylglutamylserylhistidylvalylasparaginylserylargin-368. ylmethionylglycylcysteinylglutaminylalanylthreonylglutaminylleucylthreony-369. lserylarginyltyrosylglutaminylalanylleucylleucylleucylglutaminylvalylleuc-370. ylglutamylglutaminylisoleucyllysylphenylalanylleucylglutamylglutamylgluta-371. mylisoleucylglutaminylserylleucylglutamylglutamylserylglutamylserylseryll-372. eucylserylseryltyrosylserylaspartyltryptophyltyrosylglycylserylthreonylhi-373. stidyllysylasparaginylphenylalanyllysylasparaginylvalylalanylthreonyllysy-374. lisoleucylaspartyllysylvalylaspartylthreonylvalylmethionylmethionylglycyl-375. lysyllysylleucyllysylthreonylleucylglutamylvalylleucylleucyllysylaspartyl-376. methionylglutamyllysylglycylhistidylserylleucylleucyllysylserylalanylargi-377. nylglutamyllysylglycylglutamylarginylalanylvalyllysyltyrosylleucylglutamy-378. lglutamylglycylglutamylalanylglutamylarginylleucylarginyllysylglutamyliso-379. leucylhistidylaspartylhistidylmethionylglutamylglutaminylleucyllysylgluta-380. mylleucylthreonylserylthreonylvalylarginyllysylglutamylhistidylmethionylt-381. hreonylleucylglutamyllysylglycylleucylhistidylleucylalanyllysylglutamylph-382. enylalanylserylaspartyllysylcysteinyllysylalanylleucylthreonylglutaminylt-383. ryptophylisoleucylalanylglutamyltyrosylglutaminylglutamylisoleucylleucylh-384. istidylvalylprolylglutamylglutamylprolyllysylmethionylglutamylleucyltyros-385. ylglutamyllysyllysylalanylglutaminylleucylseryllysyltyrosyllysylserylleuc-386. ylglutaminylglutaminylthreonylvalylleucylserylhistidylglutamylprolylseryl-387. valyllysylserylvalylarginylglutamyllysylglycylglutamylalanylleucylleucylg-388. lutamylleucylvalylglutaminylaspartylvalylthreonylleucyllysylaspartyllysyl-389. isoleucylaspartylglutaminylleucylglutaminylserylaspartyltyrosylglutaminyl-390. aspartylleucylcysteinylserylisoleucylglycyllysylglutamylhistidylvalylphen-391. ylalanylserylleucylglutamylalanyllysylvalyllysylaspartylhistidylglutamyla-392. spartyltyrosylasparaginylserylglutamylleucylglutaminylglutamylvalylglutam-393. yllysyltryptophylleucylleucylglutaminylmethionylserylglycylarginylleucylv-394. alylalanylprolylaspartylleucylleucylglutamylthreonylserylserylleucylgluta-395. mylthreonylisoleucylthreonylglutaminylglutaminylleucylalanylhistidylhisti-396. dyllysylalanylmethionylmethionylglutamylglutamylisoleucylalanylglycylphen-397. ylalanylglutamylaspartylarginylleucylasparaginylasparaginylleucylglutamin-398. ylmethionyllysylglycylaspartylthreonylleucylisoleucylglycylglutaminylcyst-399. einylalanylaspartylhistidylleucylglutaminylalanyllysylleucyllysylglutamin-400. ylasparaginylvalylhistidylalanylhistidylleucylglutaminylglycylthreonyllys-401. ylaspartylseryltyrosylserylalanylisoleucylcysteinylserylthreonylalanylglu-402. taminylarginylmethionyltyrosylglutaminylserylleucylglutamylhistidylglutam-403. ylleucylglutaminyllysylhistidylvalylserylarginylglutaminylaspartylthreony-404. lleucylglutaminylglutaminylcysteinylglutaminylalanyltryptophylleucylseryl-405. alanylvalylglutaminylprolylaspartylleucylglutamylprolylserylprolylglutami-。
世界上最长的单词有哪些很多人看到一个个长长的英文单词就头痛，那世界上最长的单词是什么你知道吗?下面就由店铺为你介绍下世界上最长的单词。
世界上最长单词是哪个：TITIN(肌联蛋白)的化学名称最长的英文单词，这似乎是个争论不休的议题，可见单字的组成并不如想像中的简单。
英文可以依照造字规则建立新单字，所以常常有新的长单字建立;地名是其中一个，科技词汇也很长。
我们很难去界定它。
(注意：单字的长度指的是有几个字母，而非发音或音节的长度。
)来自美国科技博客Gizmodo报道，世界上最长的单词是TITIN(肌联蛋白)的化学名称。
TITIN是一种巨型蛋白质，负责肌肉被动弹性。
它有189819个字母，全部读完需要花费约3.5小时。
世界上最长单词是哪个：各个字母开头A: antidisestablishmentarianism n. 反对教会与国家分开学说共28个字母B: biotransformation n. [生物] 生物转化共17个字母C: chemiluminescence n. [化学] 化合光;化学反光，在化学反应中，低温反光共17个字母D: dihydrostreptomycin n. [药学] 二氢链霉素(结核病特效药) 共19个字母E: electroencephalography n. [医学] 脑波纪录仪共22个字母F: flibbertigibbet n. 轻浮的女子共15个字母G: great-granddaughter n. 曾孙女;外曾孙女共18个字母H: honorificabilitudinitatibus n.不胜光荣共28个字母I: internationalization n. 国际化;国际管理化共20个字母international 国际共13个单词J: Johannisberger n.一种白葡萄酒共14个字母K: Kaiserslautern n. [地名] 凯撒劳坦(德国西部一城市) 共14个字母L: lexicostatistics n. [语言学] 词汇统计共16个字母M: microminiaturization n. [电子学] (电子设备的)超小型化共20个字母N: noncontradiction n. 无矛盾共16个字母O: omnidirectional adj. [电学] 全方向的共15个字母P: photophosphoralation [生物学] 光合磷酸化共20个字母Q: quadricentennial n. 四百年纪念(日);四百年祭典共16个字母R: radiobroadcasting n. [电子学] 无线电广播共18个字母S: self-actualization n. [心理学] 自我实现(无连词符号最长单词(血压计)：sphygmomanometer 共16个单词) 共17个字母T: tetrachloroethylene n. [化学] 四氯乙烯共19个字母U: unenthusiastically adv. 不热心地;不关心地;冷淡地共18个字母V: vasoconstriction n. [医学] 血管紧缩;血管缩小共16个字母W: well-proportioned adj. 很平衡的;(身体)很均匀的共16个字母X: xenotransplantation n. [生理]异种(生物)器官(向人体之)移植共19个字母Y: yesterdayness n. 最近;昨天性共13个字母Z: Zoroastrianism n. 袄教，拜火教共14个字母世界上最长的单词：地名最长来介绍一个稀奇古怪的地名，据说最长的是：在美国马萨诸塞州伍斯特县南部与康涅狄格州的接界处，有一个长不过4.8公里的小湖，它在地理上并无值得称道的特色，但是湖名却长得出奇，用英文表示共有44个字母：Chargoggagoggmanhaugagoggchaubunagungamaugg。
扩张性心肌病相关基因的研究进展苏晓慧;王盼;谭武红【摘要】扩张性心肌病是一种主要累及心肌的原因不明的疾病,其发病机制可能与病毒感染、免疫调节异常、遗传因素等病因有关,近年来国内外发病率均有逐年升高的趋势,预后极差.已有研究表明,可以通过基因筛查选择易感基因进行干预来治疗此病,本文对近些年已研究过的与扩张性心肌病发病机制相关的基因进行综述,为从基因水平寻找其病因及发病机制提供依据.【期刊名称】《国外医学（医学地理分册）》【年(卷),期】2014(035)001【总页数】6页(P52-56,68)【关键词】扩张性心肌病;基因;线粒体【作者】苏晓慧;王盼;谭武红【作者单位】西安交通大学医学院微量元素与地方病卫生部重点实验室,环境与疾病相关基因教育部重点实验室,陕西西安,710061;西安交通大学医学院微量元素与地方病卫生部重点实验室,环境与疾病相关基因教育部重点实验室,陕西西安,710061;西安交通大学医学院微量元素与地方病卫生部重点实验室,环境与疾病相关基因教育部重点实验室,陕西西安,710061【正文语种】中文【中图分类】R542.2扩张性心肌病(dilated cardiomyopathy DCM)是一种原因不明的心肌病，主要特征是左心室或右心室扩大、心肌收缩功能减退，常伴有心律失常，猝死率高。
近年来国内外发病率均有逐年升高的趋势,预后极差。
本文对近些年已研究过的与扩张性心肌病发病机制相关的基因进行综述，为从基因水平寻找其病因及发病机制提供依据。
扩张性心肌病的发病机制目前尚不清楚，可能与病毒感染、免疫调节异常、遗传因素等病因有关，其主要特征是单侧或双侧心腔扩大，心肌收缩功能减退。
临床表现为进行性心力衰竭、心律失常、血栓栓塞和猝死。
DCM的病理改变以心腔扩张为主，肉眼可见心肌色泽苍白、松弛，心脏扩大，心室扩大比心房明显，左心室尤为显著。
光镜下，病变心肌以灶性纤维化为主，偶有小块坏死。
心肌细胞有不同程度的肥大、萎缩、空泡变性及嗜碱变性，肌细胞间纤维组织增厚，血管及心肌细胞周围有广泛的、大小不等的纤维化病灶和淋巴细胞浸润。
1例可疑迟发性胫骨肌萎缩症的诊断分析郑光莉1，2，许叶1，段洋洋1，王埮11 海南医学院第一附属医院（第一临床学院）神经内科，海口570102；2 海南医学院摘要：目的 总结分析1例可疑迟发性胫骨肌萎缩症（TMD）患者的诊断经验。
方法 回顾性分析1例可疑迟发性TMD患者的临床资料，分析其临床症状、实验室检查、影像学检查及基因检测结果。
结果 患者女，43岁，因“四肢乏力20余年，加重2年”入院。
对背侧骨间肌、三角肌、胸锁乳突肌、股四头肌内侧头进行肌电图检查显示，未观察到自发去神经电位，插入电活动正常，肌膜稳定；轻收缩时胸锁乳突肌、股四头肌内侧头出现多相电位，右侧三角肌的运动单位缩小，可见早期的募集现象。
双侧臀部、大腿、小腿横断层面肌肉MRI检查显示，大腿肌前外侧群、后群、内侧群、盆带肌外侧群以及比目鱼肌肌肉组织几乎完全被脂肪取代，脂肪化呈网格状，或融合成斑片状，肌纤维形态消失。
肌联蛋白（TTN）基因检测结果显示，患者及其儿子存在1个杂合变异体，即c.88009+4_88009+5delins‐GT。
结合临床表现和基因检测结果以及患者家族史，考虑本例为成人起病的迟发性TMD。
患者行肌肉磁共振检查未见胫骨肌明显萎缩，建议行肌肉活检进一步明确诊断，但患者及其家属拒绝，因此未能明确诊断。
结论 迟发性TMD是一种迟发的常染色体显性远端肌病，病情进展缓慢，主要累及小腿前部肌肉，临床上可结合患者的症状、TTN 基因检测结果及肌肉活检进行诊断。
关键词：迟发性胫骨肌萎缩症；肌联蛋白基因；肌营养不良doi：10.3969/j.issn.1002-266X.2024.11.015中图分类号：R746 文献标志码：A 文章编号：1002-266X（2024）11-0067-03肌联蛋白（TTN）基因具有363个外显子，编码目前自然界中已知的最长多肽，并经过修饰后形成多种不同构象的蛋白质，常于心肌和骨骼肌中表达［1-2］。
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ucylphenylalanylalanylmethionylthreonylthr eonylglutaminylarginyltyrosylglutamylserylleucylphenylalany lalanylglutaminylleucyllysylglutamylarginy llysylglutamylglycylalanylphenylalanylvalylprolylphenylalanylvalylthreonylleucylglycylaspartylprolylglyc ylisoleucylglutamylglutaminylserylleucyllysylisoleucylaspar tylthreonylleucylisoleucylglutamylalanylglyc ylalanylaspartylalanylleucylglutamylleucylglycylisoleucylpr olylphenylalanylserylaspartylprolylleucylala nylaspartylglycylprolylthreonylisoleucylglutaminylasparagin ylalanylthreonylleucylarginylalanylphenylal anylalanylalanylglycylvalylthreonylprolylalanylglutaminylcy steinylphenylalanylglutamylmethionylleucyl alanylleucylisoleucylarginylglutaminyllysylhistidylprolylth reonylisoleucylprolylisoleucylglycylleucylleucy lmethionyltyrosylalanylasparaginylleucylvalylphenylalanylas paraginyllysylglycylisoleucylaspartylgluta mylphenylalanyltyrosylalanylglutaminylcysteinylglutamyllysy lvalylglycylvalylaspartylserylvalylleucylval ylalanylaspartylvalylprolylvalylglutaminylglutamylserylalan ylprolylphenylalanylarginylglutaminylalanyla lanylleucylarginylhistidylasparaginylvalylalanylprolylisole ucylphenylalanylisoleucylcysteinylprolylprolyl aspartylalanylaspartylaspartylaspartylleucylleucylarginylgl utaminylisoleucylalanylseryltyrosylglycylargi nylglycyltyrosylthreonyltyrosylleucylleucylserylarginylalan ylglycylvalylthreonylglycylalanylglutamylasp araginylarginylalanylalanylleucylprolylleucylasparaginylhis tidylleucylvalylalanyllysylleucyllysylglutamyl tyrosylasparaginylalanylalanylprolylprolylleucylglutaminylg lycylphenylalanylglycylisoleucylserylalanylp rolylaspartylglutaminylvalyllysylalanylalanylisoleucylaspar tylalanylglycylalanylalanylglycylalanylisoleu cylserylglycylserylalanylisoleucylvalyllysylisoleucylisoleu cylglutamylglutaminylhistidylasparaginylisoleucylglutamylprolylglutamyllysylmethionylleucylalanylalanyll eucyllysylvalylphenylalanylvalylglutaminylp rolylmethionyllysylalanylalanylthreonylarginylacetylserylty rosylserylisoleucylthreonylserylprolylserylglu taminylphenylalanylvalylphenylalanylleucylserylserylvalyltr yptophylalanylaspartylprolylisoleucylglutam ylleucylleucylasparaginylvalylcysteinylthreonylserylserylle ucylglycylasparaginylglutaminylphenylalanyl glutaminylthreonylglutaminylglutaminylalanylarginylthreonyl threonylglutaminylvalylglutaminylglutamin ylphenylalanylserylglutaminylvalyltryptophyllysylprolylphen ylalanylprolylglutaminylserylthreonylvalylar ginylphenylalanylprolylglycylaspartylvalyltyrosyllysylvalyl tyrosylarginyltyrosylasparaginylalanylvalylleu cylaspartylprolylleucylisoleucylthreonylalanylleucylleucylg lycylthreonylphenylalanylaspartylthreonylar ginylasparaginylarginylisoleucylisoleucylglutamylvalylgluta mylasparaginylglutaminylglutaminylserylpr olylthreonylthreonylalanylglutamylthreonylleucylaspartylala nylthreonylarginylarginylvalylaspartylaspar tylalanylthreonylvalylalanylisoleucylarginylserylalanylaspa raginylisoleucylasparaginylleucylvalylaspar aginylglutamylleucylvalylarginylglycylthreonylglycylleucylt yrosylasparaginylglutaminylasparaginylthreo nylphenylalanylglutamylserylmethionylserylglycylleucylvalyl tryptophylthreonylserylalanylprolylalanyltiti nmethionylglutaminylarginyltyrosylglutamylserylleucylphenyl alanylalanylisoleucylcysteinylprolylprolyla spartylalanylaspartylaspartylaspartylleucylleucylarginylglu taminylisoleucylalanylseryltyrosylglycylargin ylglycyltyrosylthreonyltyrosylleucylleucylserylarginylalanylglycylvalylthreonylglycylalanylglutamylaspar aginylarginylalanylalanylleucylprolylleucylasparaginylhisti dylleucylvalylalanyllysylleucyllysylglutamylty rosylasparaginylalanylalanylprolylprolylleucylglutaminylgly cylphenylalanylglycylisoleucylserylalanylpro lylaspartylglutaminylvalyllysylalanylalanylisoleucylasparty lalanylglycylalanylalanylglycylalanylisoleucyl serylglycylserylalanylisoleucylvalyllysylisoleucylisoleucyl glutamylglutaminylhistidylasparaginylisoleucy lglutamylprolylglutamyllysylmethionylleucylalanylalanylleuc yllysylvalylphenylalanylvalylglutaminylproly lmethionyllysylalanylalanylthreonylarginylacetylseryltyrosy lserylisoleucylthreonylserylprolylserylgluta minylphenylalanylvalylphenylalanylleucylserylserylvalyltryp tophylalanylaspartylprolylisoleucylglutamyll eucylleucylasparaginylvalylcysteinylthreonylserylserylleucy lglycylasparaginylglutaminylphenylalanylgl utaminylthreonylglutaminylglutaminylalanylarginylthreonylth reonylglutaminylvalylglutaminylglutaminyl phenylalanylserylglutaminylvalyltryptophyllysylprolylphenyl alanylprolylglutaminylserylthreonylvalylargi nylphenylalanylprolylglycylaspartylvalyltyrosyllysylvalylty rosylarginyltyrosylasparaginylalanylvalylleucy laspartylprolylleucylisoleucylthreonylalanylleucylleucylgly cylthreonylphenylalanylaspartylthreonylargin ylasparaginylarginylisoleucylisoleucylglutamylvalylglutamyl asparaginylglutaminylglutaminylserylprolylt hreonylthreonylalanylglutamylthreonylleucylaspartylalanylth reonylarginylarginylvalylaspartylaspartylal anylthreonylvalylalanylisoleucylarginylserylalanylasparagin ylisoleucylasparaginylleucylvalylasparaginylglutamylleucylvalylarginylglycylthreonylglycylleucyltyros ylasparaginylglutaminylasparaginylthreonylp henylalanylglutamylserylmethionylserylglycylleucylvalyltryp tophylthreonylserylalanylprolylalanyltitinme thionylglutaminylarginyltyrosylglutamylserylleucylphenylala nylalanylisoleucylcysteinylprolylprolylaspar tylalanylaspartylaspartylaspartylleucylleucylarginylglutami nylisoleucylalanylseryltyrosylglycylarginylgly cyltyrosylthreonyltyrosylleucylleucylserylarginylalanylglyc ylvalylthreonylglycylalanylglutamylasparagin ylarginylalanylalanylleucylprolylleucylasparaginylhistidyll eucylvalylalanyllysylleucyllysylglutamyltyrosyl asparaginylalanylalanylprolylprolylleucylglutaminylglycylph enylalanylglycylisoleucylserylalanylprolylas partylglutaminylvalyllysylalanylalanylisoleucylaspartylalan ylglycylalanylalanylglycylalanylisoleucylseryl glycylserylalanylisoleucylvalyllysylisoleucylisoleucylgluta mylglutaminylhistidylasparaginylisoleucylglut amylprolylglutamyllysylmethionylleucylalanylalanylleucyllys ylvalylphenylalanylvalylglutaminylprolylmet hionyllysylalanylalanylthreonylarginylacetylseryltyrosylser ylisoleucylthreonylserylprolylserylglutaminyl phenylalanylvalylphenylalanylleucylserylserylvalyltryptophy lalanylaspartylprolylisoleucylglutamylleucyl leucylasparaginylvalylcysteinylthreonylserylserylleucylglyc ylasparaginylglutaminylphenylalanylglutami nylthreonylglutaminylglutaminylalanylarginylthreonylthreony lglutaminylvalylglutaminylglutaminylpheny lalanylserylglutaminylvalyltryptophyllysylprolylphenylalany lprolylglutaminylserylthreonylvalylarginylphe nylalanylprolylglycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylasparaginylalanylvalylleucylaspar tylprolylleucylisoleucylthreonylalanylleucylleucylglycylthr eonylphenylalanylaspartylthreonylarginylaspa raginylarginylisoleucylisoleucylglutamylvalylglutamylaspara ginylglutaminylglutaminylserylprolylthreon ylthreonylalanylglutamylthreonylleucylaspartylalanylthreony larginylarginylvalylaspartylaspartylalanylth reonylvalylalanylisoleucylarginylserylalanylasparaginylisol eucylasparaginylleucylvalylasparaginylgluta mylleucylvalylarginylglycylthreonylglycylleucyltyrosylaspar aginylglutaminylasparaginylthreonylphenyla lanylglutamylserylmethionylserylglycylleucylvalyltryptophyl threonylserylalanylprolylalanyltitinmethionyl glutaminylarginyltyrosylglutamylserylleucylphenylalanylalan ylisoleucylcysteinylprolylprolylaspartylalan ylaspartylaspartylaspartylleucylleucylarginylglutaminylisol eucylalanylseryltyrosylglycylarginylglycyltyro sylthreonyltyrosylleucylleucylserylarginylalanylglycylvalyl threonylglycylalanylglutamylasparaginylargin ylalanylalanylleucylprolylleucylasparaginylhistidylleucylva lylalanyllysylleucyllysylglutamyltyrosylaspar aginylalanylalanylprolylprolylleucylglutaminylglycylphenyla lanylglycylisoleucylserylalanylprolylaspartyl glutaminylvalyllysylalanylalanylisoleucylaspartylalanylglyc ylalanylalanylglycylalanylisoleucylserylglycy lserylalanylisoleucylvalyllysylisoleucylisoleucylglutamylgl utaminylhistidylasparaginylisoleucylglutamyl prolylglutamyllysylmethionylleucylalanylalanylleucyllysylva lylphenylalanylvalylglutaminylprolylmethion yllysylalanylalanylthreonylarginylacetylseryltyrosylserylis oleucylthreonylserylprolylserylglutaminylphenylalanylvalylphenylalanylleucylserylserylvalyltryptophylala nylaspartylprolylisoleucylglutamylleucylleuc ylasparaginylvalylcysteinylthreonylserylserylleucylglycylas paraginylglutaminylphenylalanylglutaminylt hreonylglutaminylglutaminylalanylarginylthreonylthreonylglu taminylvalylglutaminylglutaminylphenylala nylserylglutaminylvalyltryptophyllysylprolylphenylalanylpro lylglutaminylserylthreonylvalylarginylphenyl alanylprolylglycylaspartylvalyltyrosyllysylvalyltyrosylargi nyltyrosylasparaginylalanylvalylleucylaspartyl prolylleucylisoleucylthreonylalanylleucylleucylglycylthreon ylphenylalanylaspartylthreonylarginylaspara ginylarginylisoleucylisoleucylglutamylvalylglutamylasparagi nylglutaminylglutaminylserylprolylthreonylt hreonylalanylglutamylthreonylleucylaspartylalanylthreonylar ginylarginylvalylaspartylaspartylalanylthre onylvalylalanylisoleucylarginylserylalanylasparaginylisoleu cylasparaginylleucylvalylasparaginylglutam ylleucylvalylarginylglycylthreonylglycylleucyltyrosylaspara ginylglutaminylasparaginylthreonylphenylala nylglutamylserylmethionylserylglycylleucylvalyltryptophylth reonylserylalanylprolylalanyltitinmethionylgl utaminylarginyltyrosylglutamylserylleucylphenylalanylalanyl methionylthreonylthreonylglutaminylarginy ltyrosylglutamylserylleucylphenylalanylalanylglutaminylleuc yllysylglutamylarginyllysylglutamylglycylala nylphenylalanylvalylprolylphenylalanylvalylthreonylleucylgl ycylaspartylprolylglycylisoleucylglutamylglu taminylserylleucyllysylisoleucylaspartylthreonylleucylisole ucylglutamylalanylglycylalanylaspartylalanyll eucylglutamylleucylglycylisoleucylprolylphenylalanylserylaspartylprolylleucylalanylaspartylglycylprolylt hreonylisoleucylglutaminylasparaginylalanylthreonylleucylar ginylalanylphenylalanylalanylalanylglycylv alylthreonylprolylalanylglutaminylcysteinylphenylalanylglut amylmethionylleucylalanylleucylisoleucylar ginylglutaminyllysylhistidylprolylthreonylisoleucylprolylis oleucylglycylleucylleucylmethionyltyrosylalany lasparaginylleucylvalylphenylalanylasparaginyllysylglycylis oleucylaspartylglutamylphenylalanyltyrosyl alanylglutaminylcysteinylglutamyllysylvalylglycylvalylaspar tylserylvalylleucylvalylalanylaspartylvalylpro lylvalylglutaminylglutamylserylalanylprolylphenylalanylargi nylglutaminylalanylalanylleucylarginylhistidy lasparaginylvalylalanylprolylisoleucylphenylalanylisoleucyl cysteinylprolylprolylaspartylalanylaspartylas partylaspartylleucylleucylarginylglutaminylisoleucylalanyls eryltyrosylglycylarginylglycyltyrosylthreonylt yrosylleucylleucylserylarginylalanylglycylvalylthreonylglyc ylalanylglutamylasparaginylarginylalanylalan ylleucylprolylleucylasparaginylhistidylleucylvalylalanyllys ylleucyllysylglutamyltyrosylasparaginylalanyl alanylprolylprolylleucylglutaminylglycylphenylalanylglycyli soleucylserylalanylprolylaspartylglutaminylv alyllysylalanylalanylisoleucylaspartylalanylglycylalanylala nylglycylalanylisoleucylserylglycylserylalanyli soleucylvalyllysylisoleucylisoleucylglutamylglutaminylhisti dylasparaginylisoleucylglutamylprolylglutam yllysylmethionylleucylalanylalanylleucyllysylvalylphenylala nylvalylglutaminylprolylmethionyllysylalanyl alanylthreonylarginylacetylseryltyrosylserylisoleucylthreon ylserylprolylserylglutaminylphenylalanylvalylphenylalanylleucylserylserylvalyltryptophylalanylaspartylp rolylisoleucylglutamylleucylleucylasparagin ylvalylcysteinylthreonylserylserylleucylglycylasparaginylgl utaminylphenylalanylglutaminylthreonylgluta minylglutaminylalanylarginylthreonylthreonylglutaminylvalyl glutaminylglutaminylphenylalanylserylgluta minylvalyltryptophyllysylprolylphenylalanylprolylglutaminyl serylthreonylvalylarginylphenylalanylprolylgl ycylaspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylaspa raginylalanylvalylleucylaspartylprolylleucyliso leucylthreonylalanylleucylleucylglycylthreonylphenylalanyla spartylthreonylarginylasparaginylarginyliso leucylisoleucylglutamylvalylglutamylasparaginylglutaminylgl utaminylserylprolylthreonylthreonylalanylg lutamylthreonylleucylaspartylalanylthreonylarginylarginylva lylaspartylaspartylalanylthreonylvalylalanyli soleucylarginylserylalanylasparaginylisoleucylasparaginylle ucylvalylasparaginylglutamylleucylvalylarg inylglycylthreonylglycylleucyltyrosylasparaginylglutaminyla sparaginylthreonylphenylalanylglutamylser ylmethionylserylglycylleucylvalyltryptophylthreonylserylala nylprolylalanyltitinmethionylglutaminylarginy ltyrosylglutamylserylleucylphenylalanylalanylisoleucylcyste inylprolylprolylaspartylalanylaspartylaspart ylaspartylleucylleucylarginylglutaminylisoleucylalanylseryl tyrosylglycylarginylglycyltyrosylthreonyltyros ylleucylleucylserylarginylalanylglycylvalylthreonylglycylal anylglutamylasparaginylarginylalanylalanylle ucylprolylleucylasparaginylhistidylleucylvalylalanyllysylle ucyllysylglutamyltyrosylasparaginylalanylalan ylprolylprolylleucylglutaminylglycylphenylalanylglycylisoleucylserylalanylprolylaspartylglutaminylvalylly sylalanylalanylisoleucylaspartylalanylglycylalanylalanylgly cylalanylisoleucylserylglycylserylalanylisole ucylvalyllysylisoleucylisoleucylglutamylglutaminylhistidyla sparaginylisoleucylglutamylprolylglutamyllys ylmethionylleucylalanylalanylleucyllysylvalylphenylalanylva lylglutaminylprolylmethionyllysylalanylalan ylthreonylarginylacetylseryltyrosylserylisoleucylthreonylse rylprolylserylglutaminylphenylalanylvalylphe nylalanylleucylserylserylvalyltryptophylalanylaspartylproly lisoleucylglutamylleucylleucylasparaginylval ylcysteinylthreonylserylserylleucylglycylasparaginylglutami nylphenylalanylglutaminylthreonylglutamin ylglutaminylalanylarginylthreonylthreonylglutaminylvalylglu taminylglutaminylphenylalanylserylglutamin ylvalyltryptophyllysylprolylphenylalanylprolylglutaminylser ylthreonylvalylarginylphenylalanylprolylglycy laspartylvalyltyrosyllysylvalyltyrosylarginyltyrosylasparag inylalanylvalylleucylaspartylprolylleucylisoleu cylthreonylalanylleucylleucylglycylthreonylphenylalanylaspa rtylthreonylarginylasparaginylarginylisoleu cylisoleucylglutamylvalylglutamylasparaginylglutaminylgluta minylserylprolylthreonylthreonylalanylglut amylthreonylleucylaspartylalanylthreonylarginylarginylvalyl aspartylaspartylalanylthreonylvalylalanyliso leucylarginylserylalanylasparaginylisoleucylasparaginylleuc ylvalylasparaginylglutamylleucylvalylargin ylglycylthreonylglycylleucyltyrosylasparaginylglutaminylasp araginylthreonylphenylalanylglutamylseryl methionylserylglycylleucylvalyltryptophylthreonylserylalany lprolylalanyltitinmethionylglutaminylarginyltyrosylglutamylserylleucylphenylalanylalanylisoleucylcystein ylprolylprolylaspartylalanylaspartylaspartyl aspartylleucylleucylarginylglutaminylisoleucylalanylserylty rosylglycylarginylglycyltyrosylthreonyltyrosyl leucylleucylserylarginylalanylglycylvalylthreonylglycylalan ylglutamylasparaginylarginylalanylalanylleuc ylprolylleucylasparaginylhistidylleucylvalylalanyllysylleuc yllysylglutamyltyrosylasparaginylalanylalanyl prolylprolylleucylglutaminylglycylphenylalanylglycylisoleuc ylserylalanylprolylaspartylglutaminylvalyllys ylalanylalanylisoleucylaspartylalanylglycylalanylalanylglyc ylalanylisoleucylserylglycylserylalanylisoleu cylvalyllysylisoleucylisoleucylglutamylglutaminylhistidylas paraginylisoleucylglutamylprolylglutamyllysyl methionylleucylalanylalanylleucyllysylvalylphenylalanylvaly lglutaminylprolylmethionyllysylalanylalanylt hreonylarginylacetylseryltyrosylserylisoleucylthreonylseryl prolylserylglutaminylphenylalanylvalylpheny lalanylleucylserylserylvalyltryptophylalanylaspartylprolyli soleucylglutamylleucylleucylasparaginylvalylc ysteinylthreonylserylserylleucylglycylasparaginylglutaminyl phenylalanylglutaminylthreonylglutaminylgl utaminylalanylarginylthreonylthreonylglutaminylvalylglutami nylglutaminylphenylalanylserylglutaminylv alyltryptophyllysylprolylphenylalanylprolylglutaminylserylt hreonylvalylarginylphenylalanylprolylglycylas partylvalyltyrosyllysylvalyltyrosylarginyltyrosylasparaginy lalanylvalylleucylaspartylprolylleucylisoleucylt hreonylalanylleucylleucylglycylthreonylphenylalanylaspartyl threonylarginylasparaginylarginylisoleucyli soleucylglutamylvalylglutamylasparaginylglutaminylglutaminylserylprolylthreonylthreonylalanylglutam ylthreonylleucylaspartylalanylthreonylarginylarginylvalylas partylaspartylalanylthreonylvalylalanylisoleu cylarginylserylalanylasparaginylisoleucylasparaginylleucylv alylasparaginylglutamylleucylvalylarginylgl ycylthreonylglycylleucyltyrosylasparaginylglutaminylasparag inylthreonylphenylalanylglutamylserylmet hionylserylglycylleucylvalyltryptophylthreonylserylalanylpr olylalanyltitinmethionylglutaminylarginyltyros ylglutamylserylleucylphenylalanylalanylisoleucylcysteinylpr olylprolylaspartylalanylaspartylaspartylasp artylleucylleucylarginylglutaminylisoleucylalanylseryltyros ylglycylarginylglycyltyrosylthreonyltyrosylleu cylleucylserylarginylalanylglycylvalylthreonylglycylalanylg lutamylasparaginylarginylalanylalanylleucylp rolylleucylasparaginylhistidylleucylvalylalanyllysylleucyll ysylglutamyltyrosylasparaginylalanylalanylprol ylprolylleucylglutaminylglycylphenylalanylglycylisoleucylse rylalanylprolylaspartylglutaminylvalyllysylal anylalanylisoleucylaspartylalanylglycylalanylalanylglycylal anylisoleucylserylglycylserylalanylisoleucylv alyllysylisoleucylisoleucylglutamylglutaminylhistidylaspara ginylisoleucylglutamylprolylglutamyllysylmet hionylleucylalanylalanylleucyllysylvalylphenylalanylvalylgl utaminylprolylmethionyllysylalanylalanylthre onylarginylacetylseryltyrosylserylisoleucylthreonylserylpro lylserylglutaminylphenylalanylvalylphenylala nylleucylserylserylvalyltryptophylalanylaspartylprolylisole ucylglutamylleucylleucylasparaginylvalylcyst einylthreonylserylserylleucylglycylasparaginylglutaminylphe nylalanylglutaminylthreonylglutaminylglutaminylalanylarginylthreonylthreonylglutaminylvalylglutaminyl glutaminylphenylalanylserylglutaminylvalyl tryptophyllysylprolylphenylalanylprolylglutaminylserylthreo nylvalylarginylphenylalanylprolylglycylaspar tylvalyltyrosyllysylvalyltyrosylarginyltyrosylasparaginylal anylvalylleucylaspartylprolylleucylisoleucylthre onylalanylleucylleucylglycylthreonylphenylalanylaspartylthr eonylarginylasparaginylarginylisoleucylisol eucylglutamylvalylglutamylasparaginylglutaminylglutaminylse rylprolylthreonylthr。
